Vaccinium burburan

Description: — Habit shrub or vine, epiphytic, evergreen, multi-branched. Branchlets brown, lanate when young, sparsely pubescent at maturity, with a combination of simple and stipitate-glandular trichomes, 0.3 – 3.0 mm long, rounded in cross section, 1.5 – 5.0 mm wide, lenticellate; perennating buds triangular, obtuse, 1.0 – 1.3 mm long; bud scales non-overlapping, hirsute. Leaves persistent on older branchlets, spirally and evenly arranged, condensed, overlapping, internodes 2 – 10 mm long; petiole brown, in cross section abaxially and adaxially rounded, 1.0 – 2.0 × 0.8 – 1.0 mm, pubescent; leaf blade elliptic to cordate, with larger blades on each branchlet 2.2 – 4.0 × 2.0 – 2.5 cm, coriaceous, both surfaces reddish when young turning dull green abaxially and dark green adaxially, without punctae, pubescent on both surfaces, partially glabrescent adaxially when mature, hirsute, trichomes simple, abaxial surface generally more pubescent than adaxial surface; midvein raised abaxially and adaxially, secondary veins 2 to 4 on each side of midvein with first pair arising from base and remainder along midvein, arc-ascending, abaxially raised, adaxially faintly evident or obscure, tertiary veins faintly evident or obscure, base cordate, margin entire, thinly revolute, apex acuminate, marginal glands slightly raised, ca. 1.0 mm from leaf base, one pair per leaf, 0.3 – 0.4 mm wide. Inflorescences pseudo-terminal or terminal, racemose, developing beyond confines of perennating bud, 1 per axil, 1.5 – 2.5 cm long at anthesis, 4 - or 5 - flowered; peduncle and rachis brown, terete, pubescent, trichomes same as on branchlets; bracts semi-persistent, reddish brown, non-foliaceous, ovate to elliptic, planar or occasionally cucullate, 1.0 – 1.5 × 0.5 – 0.8 mm, subcoriaceous, glabrous abaxially, pubescent adaxially, margin entire, occasionally with 1 or 2 glands near base. Pedicel ascending to slightly erect, 5.0 – 6.0 × 0.5 – 0.7 mm at anthesis, terete, pubescent; bracteoles borne at base of pedicel, linear or lanceolate, ca. 0.2 mm long, pubescent. Flowers articulated at junction with pedicel, 5.0 – 7.0 mm long. Hypanthium dark brown, triangular-obtuse, 1.0 – 1.2 × 0.8 – 1.0 mm, pubescent, trichomes same as on branchlets; calyx limb ca. 1.5 mm long; calyx lobes 5 or 6, narrowly triangular, 1.0 – 1.2 mm long, pubescent, trichomes same as on branchlets, margin entire, apex acuminate, without sessile terminal gland. Corolla narrowly conical-urceolate, tapering at middle toward apex, dark brown, 5.0 – 6.0 × 2.0 – 2.5 mm, glabrous outside except for apical portion including corolla lobes, inside glabrous; corolla lobes 5 or 6, pubescent, 0.5 – 0.7 × 0.3 – 0.4 mm, apex acute to obtuse. Stamens 8 to 10, monomorphic, distinct, 3.7 – 4.5 mm long; filaments brown, straight, dilated at base, 2.8 – 3.1 mm long, white-lanate especially toward base, trichomes 0.4 – 0.5 mm long; anthers 1.0 – 1.4 mm long, cells 0.6 – 0.9 mm long, obscurely echinulate, tubules parallel, broadly cylindrical, opening by oblique ventrally oriented apical pores, 0.4 – 0.5 mm long, pore apex shallowly crested, spurs absent. Ovary 5 - or 6 - locular but appearing pseudo- 10 - to 12 - locular with false partitions extending ca. 0.8 mm from inner wall; ovules in two columns per locule. Disk annular, bulky, conspicuously larger than hypanthium, 0.7 – 1.0 × 1.5 – 2.0 mm, puberulent, margin obscurely ridged. Style not seen. Fruit dark brown, globose, non-ridged, 3.0 – 3.5 × 2.5 – 3.0 mm, pubescent, trichomes same as on branchlets; calyx lobes persistent, slightly recurved, disk bulged, annular, ca. 1.0 × ca. 2.0 mm; fruiting pedicels ca. 5 mm long.